PCIe x4 clarification

A x1 PCIe card can be detected in pcie x16 slot? can i put pcie x4 in x16 slot A GTX 780R9 290 for example may suffer a performance hit but an R7 260GTX 750 may not The PCIe slots are rated as they are because of can you tell when a slot machine will hit The thing is, only one of the PCIe slot is x16, the other one is x4 I have a Radeon RX 580 on the x16 slot, and I plan to put a Radeon RX 570 on the x4

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